Graduate Courses

We offer a variety of courses and seminars for graduate students (there also are many other options available through other allied departments on campus. Below is a listing of most of these courses to give you a general sense of the topics that might be covered.  “Seminars” are diverse, sometimes focusing on student research, sometimes covering a topic (both from a historical and current perspective) and sometimes highlighting the most current issues in a discipline (recent examples include phenotypic plasticity, behavior genetics, biogeography, and stable isotopes). Despite this listing, your most significant growth as a scientist will often arise not by taking classes but through your individual scholarship and your informal interactions with other students and faculty (including those visiting from other institutions).


  • Ethology
  • Behavioral Ecology
  • Dynamic Modeling in Behavioral and Evolutionary Ecology
  • Mechanisms of Behavior
  • Seminar in Animal Behavior


  • Community Ecology
  • Limnology
  • Marine Communities and Oceanographic Practicum
  • Marine Ecology
  • Nutritional Ecology
  • Quantitative Methods and Ecological Inference
  • Ecological Models and Data
  • Seminar in Ecology / Frontiers and Foundations in Ecology / Readings in Population and Community Ecology
  • Tropical Biology (OTS)
  • Tropical Conservation


  • Evolutionary Ecology
  • Advanced Evolutionary Biology
  • Evolutionary Biology and Feminism
  • Evolutionary Genetics
  • Island Biogeography
  • Models and Simulations in Molecular Evolution
  • Principles of Systematic Biology
  • Research Reviews in Ecology and Evolution
  • Vertebrate Paleontology


  • Research Reviews in Ecology and Evolution
  • Biophotography
  • Graduate Orientation Seminar (required)
  • Integrative Principles (required)
  • Readings in Biology
  • Seminar in Biological Writing


  • Morphometrics
  • Reptilian Functional Morphology


  • Physiological Ecology
  • Physiological Ecology of Vertebrates
  • Seminar in Comparative Physiology


  • Avian Systematics and Biogeography
  • Concepts in Chelonian Biology
  • Ichthyology
  • Mammalian Review
  • Mammology
  • Advanced Invertebrate Zoology
  • Seminar in Herpetology